Nevertheless he that comforteth the lowly, even God, comforted us by the coming of Titus;

But God, who comforts the downcast, comforted us by the arrival of Titus,

but He who is comforting the cast-down--God--He did comfort us in the presence of Titus;

But he who encourages those that are [brought] low, [even]God, encouraged us by the coming of Titus;